Education
- Dartmouth College, Hanover, NH, A .B., 1951 Geology, Chemistry, Magna Cum Laude, Phi Beta Kappa, Rufus Choate Scholar
- Tufts University School of Medicine, Boston, MA, M.D. 1955 Medicine
- Rockefeller University, New York, NY, Ph.D., 1962 Biochemistry; Dissertation "A study of hemin biosynthesis in Haemophilus species" with Dr. Sam Granick
- Internship Hospital University of Pennsylvania, 1955-1956